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Germanus was born of prominent Romano-Gaulish parents, Rusticus and Germanilla, in Autissiodurum (Auxerre), Gaul, during the later part of the fourth century, c. 380. He received his early education in the schools of Arles and Lyons. Germanus continued his education in Rome, studying rhetoric and civil law. He practiced law in Rome before the tribunal of the prefect for some years with great success and came into contact with the court. He married Eustachia.
The emperor, Honorius, appointed him as one of the six dukes responsible for the governance of Gaul, residing in Auxerre. He led a worldly life that brought him under the displeasure of Bishop Amator of Auxerre. Angered, Germanus attempted to kill Amator, resulting in the bishop forcing him to tonsure. After ordaining him a deacon, Bp. Amator told Germanus to live as one destined to succeed him as bishop. Taking the whole incident as an action of the Holy Spirit, Germanus changed his life completely and as an ascetic he devoted himself to prayer, study, and charity.